The tragic sense becomes the keynote of his novels.

悲剧的基调。

That an acute, tragic sense of the frailty of human existence framed Turner's life, and powers the greatest of his works.

正是对人生脆弱的那种敏锐且悲伤的意识塑造了特纳的生活，也给他那些伟大的作品赋予了力量。

The tragic sense turns into despair.

悲剧性变成绝望。