While other tissue types are mainly made of living cells packed together, the inert matrix between connective-tissue cells is actually more important than what's inside the cells.

虽然其他组织类型主要由堆积在一起的活细胞组成，但结缔组织细胞之间的惰性基质实际上比细胞内部的东西更重要。

The innermost layer tends to have loosely packed cells that retain their nuclei despite being filled with keratin, a strong protein, creating what's called a " wet keratin" appearance.

最内层往往分布着排列松散的有核细胞，尽管这些细胞充满了高强度的角蛋白，形成一种称为“湿性角化”的外观。