Through the years they have made significant contributions to species conservation.多年来他们对物种保护做出了重大贡献。He is seen as having made a significant contribution towards the eradication of corruption.人们认为他已经为铲除腐败作出了重大贡献。He made many valuable contributions to the field of science.他在科学领域做出了许多重大贡献。