Logan Kistler, an archaeologist at the Smithsonian's National Museum of Natural History.

史密森尼国家自然历史博物馆的考古学家洛根·基斯特勒说到。

Kistler says mastodons probably weren't put off by the gourds' bitter taste.

基斯特勒表示，乳齿象可能不会被野生南瓜的苦味吓退。

Kistler reported those findings in the Proceedings of the National Academy of Sciences in 2015.

基斯特勒在2015年的《美国国家科学院院刊》上报告了这些发现。

And humans, Kistler says, which also tend to disturb the environments around them, creating great squash habitat—may have taken the mastodon's place.

而人类，奇斯特勒说， 它们也往往会扰乱周围的环境，创造出巨大的南瓜栖息地——可能已经取代了乳齿象的位置。