Inside the duodenal cells, iron molecules bind to a protein called ferritin, which temporarily stores the iron.

在十二指肠细胞内，含铁分子与一种称为铁蛋白的蛋白质结合，铁蛋白会暂时贮存铁。

A generous amount of bicarbonate is also secreted by a number of glands lying in the submucosa of the duodenal wall.

十二指肠肠壁上分布的大量腺体，也可分泌大量的碳酸氢盐。

Early in 1917 he began suffering from a duodenal ulcer and he would eventually have to have this operated on early that winter.

1917 年初，他开始患有十二指肠溃疡，最终不得不在那个冬初接受手术。