They are the first footprint evidence that some ancient ancestors of modern crocodiles walked on two legs.
它们是现代鳄鱼的一些远古祖先用两条腿行走的第一个足迹证据。
PBS地球-动物趣闻
Now, it can be pretty easy to figure out what made a modern trace - because we can actually observe it happening - like, watching a crocodile walk through mud, leaving its footprints behind.